City Trees

July/August 2021

City Trees is a premier publication focused on urban + community forestry. In each issue, you’ll learn how to best manage the trees in your community and more!

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Axillary Wood In all these images, I have been showing frac- ture surfaces of broken branch junctions, so some of the images in this article have included glimpses of some axillary wood, which I have cate- gorized as a reaction wood (Slater et al., 2014). As with all reaction woods, there is a drift in tissue qualities as there is a graduated transition from 'normal' wood to a reaction wood. Axillary wood is most distinctly different in its center, right in the center of the branch union: cutting further and further into a branch junction is a bit like cutting into the middle of an onion. The tissues are changing qualita- tively as you get nearer and nearer to the center. Pictured is some axillary wood of common beech when I fractured a branch junction. Probably the most important practical aspect of the twisting and turning that the wood grain of axillary wood does is that it causes some wood cells to be orientated so that you must stretch them and break them along their length to break the branch junction. This requires a great deal more energy to break this wood than if all cells were orientated radially or tangentially. Axillary wood is the latest key finding in wood anatomy, from an arboricultural perspective. It is a key responsive woody tissue that is much denser than surrounding wood, much more crack-resistant, much stronger in the radial and tangential directions—but not much use in the conduction of sap (Slater et al., 2014). As a tissue type formed in all trees which have branches, it warrants much more scientific scrutiny. It has been my pleasure to introduce its importance into arboricultural texts which have, for many years, ignored its existence or not realized its significance. That is why they awarded me a PhD—that I had iden- tified an important biomechanical role played by this key woody tissue, which helps to form the branching structure of trees. It is easy to find if you know what you are looking for, as it is the denser wood that is typically much more difficult to split with an axe, that lies in the crotch of a branch junction due to the swirling character of the wood grain in that location. The consequences of understanding the role of axil- lary wood in trees are many, so I invite readers that found this short journey to 'Planet Wood' inter- esting to follow up with some of my publications that help to define the properties of axillary wood. I wonder how many more lessons can be learnt from close examination of the wood that trees grow. 30 CityTREES

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